diff --git a/src/AutoMapper/ExpressionExtensions.cs b/src/AutoMapper/ExpressionExtensions.cs
index 7e3291c6fe..5fb6972a6b 100644
--- a/src/AutoMapper/ExpressionExtensions.cs
+++ b/src/AutoMapper/ExpressionExtensions.cs
@@ -166,7 +166,6 @@ protected override Expression VisitMember(MemberExpression node)
         internal class RemoveIfNotNullVisitor : ExpressionVisitor
         {
             private readonly Expression[] _expressions;
-            private readonly IList<Expression> AllreadyUpdated = new List<Expression>();
 
             public RemoveIfNotNullVisitor(params Expression[] expressions)
             {
@@ -196,9 +195,12 @@ public ConvertingVisitor(ParameterExpression oldParam, Expression newParam)
 
             protected override Expression VisitMember(MemberExpression node)
             {
-                return node.Expression == _oldParam
-                    ? MakeMemberAccess(ToType(_newParam, _oldParam.Type), node.Member)
-                    : base.VisitMember(node);
+                if (node.Expression == _oldParam)
+                {
+                    node = MakeMemberAccess(ToType(_newParam, _oldParam.Type), node.Member);
+                }
+
+                return base.VisitMember(node);
             }
 
             protected override Expression VisitParameter(ParameterExpression node)
@@ -208,9 +210,12 @@ protected override Expression VisitParameter(ParameterExpression node)
 
             protected override Expression VisitMethodCall(MethodCallExpression node)
             {
-                return node.Object == _oldParam
-                    ? Call(ToType(_newParam, _oldParam.Type), node.Method, node.Arguments)
-                    : base.VisitMethodCall(node);
+                if (node.Object == _oldParam)
+                {
+                    node = Call(ToType(_newParam, _oldParam.Type), node.Method, node.Arguments);
+                }
+
+                return base.VisitMethodCall(node);
             }
         }
 
@@ -227,7 +232,10 @@ public ReplaceExpressionVisitor(Expression oldExpression, Expression newExpressi
 
             public override Expression Visit(Expression node)
             {
-                return _oldExpression == node ? _newExpression : base.Visit(node);
+                if (_oldExpression == node)
+                    node = _newExpression;
+
+                return base.Visit(node);
             }
         }
 
diff --git a/src/UnitTests/Bug/MapFromClosureBug.cs b/src/UnitTests/Bug/MapFromClosureBug.cs
new file mode 100644
index 0000000000..4426b93753
--- /dev/null
+++ b/src/UnitTests/Bug/MapFromClosureBug.cs
@@ -0,0 +1,72 @@
+namespace AutoMapper.UnitTests.Bug
+{
+    using System;
+    using Rhino.Mocks;
+    using Should;
+
+    public class MapFromClosureBug : SpecBaseBase
+    {
+        private static readonly IDateProvider _dateProvider;
+
+        static MapFromClosureBug()
+        {
+            _dateProvider = MockRepository.GenerateMock<IDateProvider>();
+        }
+
+        public interface IDateProvider
+        {
+            DateTime CurrentRestaurantTime(Restaurant restaurant);
+        }
+
+        public class Result
+        {
+            public Booking Booking { get; set; }
+        }
+
+        public class Restaurant
+        {
+        }
+
+        public class Booking
+        {
+            public Restaurant Restaurant { get; set; }
+
+            public int? CalculateTotal(DateTime currentTime)
+            {
+                return null;
+            }
+        }
+
+        public class ResultDto
+        {
+            public BookingDto Booking { get; set; }
+        }
+
+        public class BookingDto
+        {
+            public int? Total { get; set; }
+        }
+
+        public void Should_map_successfully()
+        {
+            var mapperConfiguration = new MapperConfiguration(cfg =>
+            {
+                cfg.CreateMap<Result, ResultDto>();
+                cfg.CreateMap<Booking, BookingDto>()
+                    .ForMember(d => d.Total,
+                        o => o.MapFrom(b => b.CalculateTotal(_dateProvider.CurrentRestaurantTime(b.Restaurant))));
+            });
+
+            var mapper = mapperConfiguration.CreateMapper();
+
+            var result = new Result { Booking = new Booking() };
+
+            // Act
+            var dto = mapper.Map<ResultDto>(result);
+
+            // Assert
+            dto.ShouldNotBeNull();
+        }
+
+    }
+}
\ No newline at end of file
diff --git a/src/UnitTests/UnitTests.Net4.csproj b/src/UnitTests/UnitTests.Net4.csproj
index 898e559d99..2e7b212965 100644
--- a/src/UnitTests/UnitTests.Net4.csproj
+++ b/src/UnitTests/UnitTests.Net4.csproj
@@ -110,6 +110,7 @@
     <Compile Include="Bug\CollectionBaseClassGetConvention.cs" />
     <Compile Include="Bug\InitializeNRE.cs" />
     <Compile Include="Bug\GuidTryExpression.cs" />
+    <Compile Include="Bug\MapFromClosureBug.cs" />
     <Compile Include="Bug\NullToString.cs" />
     <Compile Include="Bug\NullSubstituteInnerClass.cs" />
     <Compile Include="Bug\MapExpandoObjectProperty.cs" />