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Distance dependent structural plasticity #3369

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@FinnBurkhardt FinnBurkhardt commented Nov 18, 2024

Description

This pull request introduces new functionality to handle distance-dependent connection probabilities in the SPManager class. The following methods have been added or modified:

New Features

  • gather_global_positions_and_ids()
    Collects and verifies global neuron positions and IDs.
    Ensures that neurons are spatially distributed, raising an error if positions are undefined while distance dependency is enabled.

  • gaussian_kernel()
    Computes a probability based on the Gaussian distance kernel between two neuron positions.

  • build_probability_list()
    Constructs a probability list for all neuron pairs based on their spatial positions.
    Supports caching for efficient repeated lookups.

  • get_neuron_pair_index()
    Maps two neuron IDs to a unique index for accessing pair-specific probabilities in the probability_list.

  • global_shuffle_spatial()
    Implements distance-dependent shuffling for pairing neurons based on spatial proximity and probability.

  • roulette_wheel_selection()
    Performs a weighted random selection of post-synaptic neurons based on precomputed probabilities.

Modified Methods

  • enable_structural_plasticity()
    Integrates calls to gather_global_positions_and_ids() and build_problist() when distance dependency is enabled.

Distance-Dependent Features

  • Enable Distance Dependency
    Distance dependency can be enabled by setting structural_plasticity_gaussian_kernel_sigma to a positive value. This parameter controls the width of the Gaussian kernel used for connection probabilities. Higher values result in reduced sensitivity to distance, making distant neurons more likely to connect.

  • Caching Connection Probabilities
    If structural_plasticity_cache_probabilities is set to true, connection probabilities will be precomputed and cached. This can improve performance, particularly in long-running simulations.


Testing

The following tests have been added to validate the new functionality:

C++ Tests:

  • test_gaussian_kernel
    Tests Gaussian kernel computations for various distances and sigma values.

  • test_get_neuron_pair_index
    Confirms correct mapping of neuron pairs to unique indices.

  • test_global_shuffle_spatial
    Validates spatially aware shuffling logic.
    Ensures connections favor spatial proximity and prevent self-connections.

Python Tests:

  • test_distance_dependent_without_positions()
    Ensures an error is raised when distance dependency is enabled, but positions are not provided.

  • test_gaussian_kernel()
    Tests probability computation using Gaussian distance dependency.

  • test_probability_list_building()
    Verifies the correct initialization and population of the probability list.

  • test_roulette_wheel_selection()
    Confirms that weighted random selection behaves as expected.

  • test_structural_plasticity_with_positions()
    Ensures structural plasticity works correctly when neuron positions are defined.

  • test_synapse_creation_distance_dependent()
    Validates distance-dependent synapse formation using position data.


Related Issue:

Incorporate Distance-Dependent Connection Probability in Structural Plasticity Module #3209

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